
Entwined lives by Tim Laman, US —
Category: Grand Prize An endangered young male Bornean Orangutan climbs over 30 meters up in the rain forest of Gunung Palung National Park in Indonesian Borneo. The park is one of the few protected orangutan environments in Borneo.

The alley cat by Nayan Khanolkar, India —
Category: UrbanAt night, in a suburb of Mumbai bordering Sanjay Gandhi National Park, leopards slip ghost-like through the maze of alleys, looking for food. Especially stray dogs. The big cats are largely accepted as a part of life and culture in the region, and even revered in traditional paintings displayed on the walls of local homes.

Eviction attempt by Ganesh H Shankar , India —
Category: BirdsIndian rose-ringed parakeets attack a Bengal monitor lizard that had invaded their nest, high in a tree in India's Keoladeo National Park. After two days and repeated attacks, the parakeets eventually gave up and left to find another home.

The moon and the crow by Gideon Knight, UK —
Category: Grand Prize (Young Wildlife Photographer)A crow perching in a tree is a common scene in a London park, but when set against the blue light of dusk and a full moon it felt 'almost supernatural, like something out of a fairy tale,' says Gideon.

The pangolin pit by Paul Hilton, UK/Australia —
Category: Photojournalist Award for a single imagePaul Hilton captured this scene of some 4,000 defrosting pangolins from one of the largest seizures on record of the heavily trafficked animals. The scaly mammals are prized in China and Vietnam for their exotic meat and for their scales used in traditional medicine. Four of the eight existing species are endangered as a result of the trade.

Star player by Luis Javier Sandoval, Mexico —
Category: ImpressionsA young sea lion swimming in the Gulf of California grabs a starfish, which it then offered to photographer Luis Javier Sandoval. Sandoval says, 'I love the way sea lions interact with divers and how smart they are.'

The sand canvas by Rudi Sebastian, Germany —
Category: DetailsThe pristine white sand of Brazil's Lençóis Maranhenses National Park offers a blank canvas to natural processes of rain, bacteria, streams, and evaporation that leave behind elegant patterns in the dunes, which can tower to 40 meters (130 feet) high.

Wind composition by Valter Binotto, Italy —
Category: Plants and FungiWith every gust of wind, showers of pollen are released from a hazel tree and illuminated by the winter sun near photographer Valter Binotto's home in northern Italy.

Snapper party by Tony Wu, US —
Category: UnderwaterThousands of two‑spot red snapper fish gather to spawn around Palau in the western Pacific Ocean for several days each month in sync with the full moon of the lunar cycle.

Requiem for an owl by Mats Andersson, Sweden —
Category: Black and WhiteA solitary Eurasian pygmy owl was photographed by Mats Andersson after he discovered the owl's mate lying dead on the forest floor. "The owl's resting posture reflected my sadness for its lost companion," says Andersson. These birds of prey are known to form pair bonds in autumn that last through spring, and sometimes for more than one breeding season.